Zealandia pustulata(G.Forst.) Testo & A.R.Field

hound's tongue fern

WFO wfo-1000034943 Accepted WFO 2026-06 8 photographs CC0 / CC BY

Plate 1 figs. a–h

Zealandia pustulata, photographed by Jon Sullivan
fig. a Jon Sullivan, CC BY 4.0 / 2022-05-01 / obs. 195810862
Zealandia pustulata, photographed by Halema J
fig. b Halema J, CC BY 4.0 / obs. 192756913
Zealandia pustulata, photographed by L M Maule
fig. c L M Maule, CC0 1.0 / obs. 192654618
Zealandia pustulata, photographed by Alice Shanks
fig. d Alice Shanks, CC BY 4.0 / obs. 193394805
Zealandia pustulata, photographed by Jon Sullivan
fig. e Jon Sullivan, CC BY 4.0 / obs. 195625231
Zealandia pustulata, photographed by Jon Sullivan
fig. f Jon Sullivan, CC BY 4.0 / obs. 195622134
Zealandia pustulata, photographed by Jaco Grundling
fig. g Jaco Grundling, CC0 1.0 / obs. 192112291
Zealandia pustulata, photographed by Jon Sullivan
fig. h Jon Sullivan, CC BY 4.0 / obs. 195607901

Every figure is a research-grade observation under CC0, CC BY or CC BY-SA, rehosted with the photographer’s name, the licence and the observation it came from. Photographs under a NonCommercial licence are excluded from this site and are never stored, which costs us a great many pictures and is not negotiable.

Also published as 25 synonyms

A synonym is not an error. It is a record of botanists disagreeing, in print, about where this plant belongs. Each of these was somebody’s considered answer.

  • Chrysopteris billardierei Link
  • Chrysopteris lepidopoda Link
  • Drynaria billardierei J.Sm.
  • Drynaria pustulata (G.Forst.) J.Sm.
  • Lecanopteris pustulata subsp. howensis (Tindale & P.S.Green) Perrie & Brownsey
  • Lecanopteris pustulata subsp. pustulata (G.Forst.) Perrie & Brownsey
  • Microsorum diversifolium (Willd.) Copel.
  • Microsorum pustulatum (G.Forst.) Copel.
  • Microsorum pustulatum subsp. howense (Tindale & P.S.Green) Bostock
  • Phymatodes billardierei C.Presl
  • Phymatodes diversifolia (Willd.) Pic.Serm.
  • Phymatodes pustulata (G.Forst.) C.Presl
  • Phymatosorus diversifolius (Willd.) Pic.Serm.
  • Phymatosorus pustulatus (G.Forst.) M.F.Large, Braggins & P.S.Green
  • Phymatosorus pustulatus subsp. howensis Tindale & P.S.Green
  • Pleopeltis billardierei T.Moore
  • Pleopeltis diversifolia (Willd.) Melvaine
  • Pleopeltis lepidopoda T.Moore
  • Pleopeltis pustulata (G.Forst.) T.Moore
  • Polypodium billardierei R.Br.
  • Polypodium diversifolium Willd.
  • Polypodium lepidopodum Link
  • Polypodium pustulatum G.Forst.
  • Polypodium scandens Labill.

and 1 more.

Sourcesevery claim on this page

  1. World Flora Online Plant List. accepted name, authority, classification. CC0. Retrieved 2026-07-12.
  2. iNaturalist. photographs and flowering annotations, CC0 / CC BY / CC BY-SA only. per photograph. Retrieved 2026-06-27.

We publish what we can source and we say so when we cannot. This page has no care advice, no toxicity claim and no native range, because we do not yet have those from a source we can cite.