{
  "$schema": "https://botanai.app/schema/species-v1.json",
  "schema_version": 1,
  "url": "https://botanai.app/plants/ludwigia-palustris/",
  "last_modified": "2025-12-15",
  "id": {
    "slug": "ludwigia-palustris",
    "wfo": "wfo-1000070656",
    "inat_taxon_id": 60221
  },
  "name": {
    "scientific": "Ludwigia palustris",
    "authorship": "(L.) Elliott",
    "genus": "Ludwigia",
    "epithet": "palustris",
    "family": "Onagraceae",
    "rank": "species",
    "taxonomic_status": "accepted",
    "accepted_according_to": "World Flora Online Plant List, 2026-06",
    "is_homonym": false,
    "published_in": "Sketch Bot. S. Carolina  1: 211 (1817)",
    "protologue_url": "https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:30031106-2",
    "common": {
      "preferred": "marsh seedbox",
      "english": [
        "marsh seedbox"
      ],
      "source": "usda"
    },
    "synonyms": [
      {
        "name": "Dantia palustris",
        "authorship": "Des Moul."
      },
      {
        "name": "Isnardia ascendens",
        "authorship": "Hall ex Eaton & Wright"
      },
      {
        "name": "Isnardia nitida",
        "authorship": "Poir."
      },
      {
        "name": "Isnardia palustris",
        "authorship": "L."
      },
      {
        "name": "Isnardia palustris f. submersa",
        "authorship": "Glück"
      },
      {
        "name": "Isnardia palustris var. americana",
        "authorship": "DC."
      },
      {
        "name": "Isnardia palustris var. riparia",
        "authorship": "Boenn."
      },
      {
        "name": "Isnardia palustris var. rivularis",
        "authorship": "Boenn."
      },
      {
        "name": "Jussiaea isnardia",
        "authorship": "E.H.L.Krause"
      },
      {
        "name": "Ludwigia apetala",
        "authorship": "Walter"
      },
      {
        "name": "Ludwigia nitida",
        "authorship": "Michx."
      },
      {
        "name": "Ludwigia palustris f. elongata",
        "authorship": "Fassett"
      },
      {
        "name": "Ludwigia palustris f. submersa",
        "authorship": "Eames"
      },
      {
        "name": "Ludwigia palustris var. americana",
        "authorship": "(DC.) Fernald & Griscom"
      },
      {
        "name": "Ludwigia palustris var. inundata",
        "authorship": "Svenson"
      },
      {
        "name": "Ludwigia palustris var. liebmannii",
        "authorship": "H.Lév."
      },
      {
        "name": "Ludwigia palustris var. nana",
        "authorship": "Fernald & Griscom"
      },
      {
        "name": "Ludwigia palustris var. pacifica",
        "authorship": "Fernald & Griscom"
      },
      {
        "name": "Ludwigia palustris var. typica",
        "authorship": "Fernald & Griscom"
      },
      {
        "name": "Quadricosta palustris",
        "authorship": "Dulac"
      }
    ],
    "wikipedia": null
  },
  "photographs": {
    "shown": 8,
    "usable_found": 312,
    "licences": [
      "CC0",
      "CC-BY",
      "CC-BY-SA"
    ],
    "noncommercial_excluded": true,
    "items": [
      {
        "photo_id": 196068486,
        "license": "CC-BY",
        "license_name": "CC BY 4.0",
        "license_url": "https://creativecommons.org/licenses/by/4.0/",
        "derivatives_restricted": false,
        "creator": "Jody Shugart",
        "creator_url": "https://www.inaturalist.org/people/terrapinjoe",
        "source_url": "https://www.inaturalist.org/observations/d32acac9-9bc7-4ea2-b34b-57addd271ab7",
        "observed_on": "2022-05-05",
        "lat": 30.198,
        "lon": -91.342,
        "original": {
          "width": 1024,
          "height": 768
        },
        "widths": [
          400,
          800
        ],
        "urls": {
          "avif": {
            "400": "https://img.botanai.app/p/196068486/r1/400.avif",
            "800": "https://img.botanai.app/p/196068486/r1/800.avif"
          },
          "jpeg": {
            "400": "https://img.botanai.app/p/196068486/r1/400.jpg",
            "800": "https://img.botanai.app/p/196068486/r1/800.jpg"
          }
        },
        "attribution": "Jody Shugart, CC BY 4.0, via iNaturalist"
      },
      {
        "photo_id": 195432967,
        "license": "CC-BY-SA",
        "license_name": "CC BY-SA 4.0",
        "license_url": "https://creativecommons.org/licenses/by-sa/4.0/",
        "derivatives_restricted": true,
        "creator": "Meghan Cassidy",
        "creator_url": "https://www.inaturalist.org/people/wildcarrot",
        "source_url": "https://www.inaturalist.org/observations/1e7a4285-c9c7-4845-9831-6edfc7b12e64",
        "observed_on": "2022-05-02",
        "lat": 32.966,
        "lon": -96.942,
        "original": {
          "width": 1024,
          "height": 683
        },
        "widths": [
          400,
          800
        ],
        "urls": {
          "avif": {
            "400": "https://img.botanai.app/p/195432967/r1/400.avif",
            "800": "https://img.botanai.app/p/195432967/r1/800.avif"
          },
          "jpeg": {
            "400": "https://img.botanai.app/p/195432967/r1/400.jpg",
            "800": "https://img.botanai.app/p/195432967/r1/800.jpg"
          }
        },
        "attribution": "Meghan Cassidy, CC BY-SA 4.0, via iNaturalist"
      },
      {
        "photo_id": 182787141,
        "license": "CC-BY",
        "license_name": "CC BY 4.0",
        "license_url": "https://creativecommons.org/licenses/by/4.0/",
        "derivatives_restricted": false,
        "creator": "Mark Apgar",
        "creator_url": "https://www.inaturalist.org/people/apgarm",
        "source_url": "https://www.inaturalist.org/observations/e4a5501c-6077-46f2-beda-7975fd2313c5",
        "observed_on": "2022-03-11",
        "lat": 39.382,
        "lon": -77.165,
        "original": {
          "width": 1024,
          "height": 1024
        },
        "widths": [
          400,
          800
        ],
        "urls": {
          "avif": {
            "400": "https://img.botanai.app/p/182787141/r1/400.avif",
            "800": "https://img.botanai.app/p/182787141/r1/800.avif"
          },
          "jpeg": {
            "400": "https://img.botanai.app/p/182787141/r1/400.jpg",
            "800": "https://img.botanai.app/p/182787141/r1/800.jpg"
          }
        },
        "attribution": "Mark Apgar, CC BY 4.0, via iNaturalist"
      },
      {
        "photo_id": 182184016,
        "license": "CC-BY",
        "license_name": "CC BY 4.0",
        "license_url": "https://creativecommons.org/licenses/by/4.0/",
        "derivatives_restricted": false,
        "creator": "Saryu Mae 前 朝琉",
        "creator_url": "https://www.inaturalist.org/people/invertebratist",
        "source_url": "https://www.inaturalist.org/observations/597f7f7a-8dab-46b6-80d6-be40bd014107",
        "observed_on": "2022-03-05",
        "lat": -36.878,
        "lon": 174.627,
        "original": {
          "width": 1024,
          "height": 768
        },
        "widths": [
          400,
          800
        ],
        "urls": {
          "avif": {
            "400": "https://img.botanai.app/p/182184016/r1/400.avif",
            "800": "https://img.botanai.app/p/182184016/r1/800.avif"
          },
          "jpeg": {
            "400": "https://img.botanai.app/p/182184016/r1/400.jpg",
            "800": "https://img.botanai.app/p/182184016/r1/800.jpg"
          }
        },
        "attribution": "Saryu Mae 前 朝琉, CC BY 4.0, via iNaturalist"
      },
      {
        "photo_id": 178192783,
        "license": "CC-BY",
        "license_name": "CC BY 4.0",
        "license_url": "https://creativecommons.org/licenses/by/4.0/",
        "derivatives_restricted": false,
        "creator": "Leon Perrie",
        "creator_url": "https://www.inaturalist.org/people/leonperrie",
        "source_url": "https://www.inaturalist.org/observations/d512962b-93e5-45bc-b418-534897074e8d",
        "observed_on": "2022-02-04",
        "lat": -40.518,
        "lon": 172.743,
        "original": {
          "width": 1024,
          "height": 768
        },
        "widths": [
          400,
          800
        ],
        "urls": {
          "avif": {
            "400": "https://img.botanai.app/p/178192783/r1/400.avif",
            "800": "https://img.botanai.app/p/178192783/r1/800.avif"
          },
          "jpeg": {
            "400": "https://img.botanai.app/p/178192783/r1/400.jpg",
            "800": "https://img.botanai.app/p/178192783/r1/800.jpg"
          }
        },
        "attribution": "Leon Perrie, CC BY 4.0, via iNaturalist"
      },
      {
        "photo_id": 176107790,
        "license": "CC-BY",
        "license_name": "CC BY 4.0",
        "license_url": "https://creativecommons.org/licenses/by/4.0/",
        "derivatives_restricted": false,
        "creator": "naturewatchwidow",
        "creator_url": "https://www.inaturalist.org/people/naturewatchwidow",
        "source_url": "https://www.inaturalist.org/observations/61964a3f-d662-4a76-b21f-92940cbd17a2",
        "observed_on": "2022-01-16",
        "lat": -41.255,
        "lon": 175.176,
        "original": {
          "width": 1024,
          "height": 768
        },
        "widths": [
          400,
          800
        ],
        "urls": {
          "avif": {
            "400": "https://img.botanai.app/p/176107790/r1/400.avif",
            "800": "https://img.botanai.app/p/176107790/r1/800.avif"
          },
          "jpeg": {
            "400": "https://img.botanai.app/p/176107790/r1/400.jpg",
            "800": "https://img.botanai.app/p/176107790/r1/800.jpg"
          }
        },
        "attribution": "naturewatchwidow, CC BY 4.0, via iNaturalist"
      },
      {
        "photo_id": 172652823,
        "license": "CC0",
        "license_name": "CC0 1.0",
        "license_url": "https://creativecommons.org/publicdomain/zero/1.0/",
        "derivatives_restricted": false,
        "creator": "kcthetc1",
        "creator_url": "https://www.inaturalist.org/people/kcthetc1",
        "source_url": "https://www.inaturalist.org/observations/a0290fb3-33ad-49e9-8d07-b2ce17981c92",
        "observed_on": "2021-12-15",
        "lat": 32.303,
        "lon": -80.967,
        "original": {
          "width": 1024,
          "height": 768
        },
        "widths": [
          400,
          800
        ],
        "urls": {
          "avif": {
            "400": "https://img.botanai.app/p/172652823/r1/400.avif",
            "800": "https://img.botanai.app/p/172652823/r1/800.avif"
          },
          "jpeg": {
            "400": "https://img.botanai.app/p/172652823/r1/400.jpg",
            "800": "https://img.botanai.app/p/172652823/r1/800.jpg"
          }
        },
        "attribution": "kcthetc1, CC0 1.0, via iNaturalist"
      },
      {
        "photo_id": 167881850,
        "license": "CC-BY",
        "license_name": "CC BY 4.0",
        "license_url": "https://creativecommons.org/licenses/by/4.0/",
        "derivatives_restricted": false,
        "creator": "Kevin Faccenda",
        "creator_url": "https://www.inaturalist.org/people/kevinfaccenda",
        "source_url": "https://www.inaturalist.org/observations/4800257a-081f-417d-8f43-c5037e52b966",
        "observed_on": "2021-10-31",
        "lat": 21.549,
        "lon": -157.883,
        "original": {
          "width": 1024,
          "height": 768
        },
        "widths": [
          400,
          800
        ],
        "urls": {
          "avif": {
            "400": "https://img.botanai.app/p/167881850/r1/400.avif",
            "800": "https://img.botanai.app/p/167881850/r1/800.avif"
          },
          "jpeg": {
            "400": "https://img.botanai.app/p/167881850/r1/400.jpg",
            "800": "https://img.botanai.app/p/167881850/r1/800.jpg"
          }
        },
        "attribution": "Kevin Faccenda, CC BY 4.0, via iNaturalist"
      }
    ]
  },
  "native_range": {
    "scheme": "TDWG World Geographical Scheme for Recording Plant Distributions, level 3",
    "source": "Kew, World Checklist of Vascular Plants (WCVP v16)",
    "license": "CC BY 3.0",
    "excludes": [
      "introduced",
      "extinct",
      "location_doubtful"
    ],
    "codes": [
      "ALG",
      "ANG",
      "BOT",
      "CPP",
      "ZAI",
      "NAT",
      "LES",
      "MOR",
      "NAM",
      "TVL",
      "TUN",
      "ZAM",
      "ZIM",
      "IRN",
      "LBS",
      "NCS",
      "TCS",
      "TUR",
      "ALB",
      "AUT",
      "BGM",
      "BUL",
      "COR",
      "CZE",
      "FRA",
      "GER",
      "GRB",
      "GRC",
      "HUN",
      "ITA",
      "YUG",
      "NET",
      "POL",
      "POR",
      "ROM",
      "SAR",
      "SPA",
      "SWI",
      "TUE",
      "UKR",
      "ALA",
      "ARI",
      "ARK",
      "BRC",
      "CAL",
      "CNT",
      "DEL",
      "WDC",
      "FLA",
      "GEO",
      "IDA",
      "ILL",
      "INI",
      "IOW",
      "KAN",
      "KTY",
      "LOU",
      "MAI",
      "MRY",
      "MAS",
      "MXC",
      "MXG",
      "MXE",
      "MXN",
      "MXT",
      "MXS",
      "MIC",
      "MIN",
      "MSI",
      "MSO",
      "NEB",
      "NBR",
      "NWH",
      "NWJ",
      "NWM",
      "NWY",
      "NFL",
      "NCA",
      "NSC",
      "OHI",
      "OKL",
      "ONT",
      "ORE",
      "PEN",
      "QUE",
      "RHO",
      "SCA",
      "TEN",
      "TEX",
      "VER",
      "VRG",
      "WAS",
      "WVA",
      "WIS",
      "CLM",
      "COS",
      "CUB",
      "DOM",
      "GUA",
      "HAI",
      "JAM",
      "PUE",
      "VEN"
    ],
    "regions": [
      {
        "code": "ALG",
        "name": "Algeria",
        "continent": "AFRICA",
        "region": "Northern Africa"
      },
      {
        "code": "ANG",
        "name": "Angola",
        "continent": "AFRICA",
        "region": "South Tropical Africa"
      },
      {
        "code": "BOT",
        "name": "Botswana",
        "continent": "AFRICA",
        "region": "Southern Africa"
      },
      {
        "code": "CPP",
        "name": "Cape Provinces",
        "continent": "AFRICA",
        "region": "Southern Africa"
      },
      {
        "code": "ZAI",
        "name": "DR Congo",
        "continent": "AFRICA",
        "region": "West-Central Tropical Africa"
      },
      {
        "code": "NAT",
        "name": "KwaZulu-Natal",
        "continent": "AFRICA",
        "region": "Southern Africa"
      },
      {
        "code": "LES",
        "name": "Lesotho",
        "continent": "AFRICA",
        "region": "Southern Africa"
      },
      {
        "code": "MOR",
        "name": "Morocco",
        "continent": "AFRICA",
        "region": "Northern Africa"
      },
      {
        "code": "NAM",
        "name": "Namibia",
        "continent": "AFRICA",
        "region": "Southern Africa"
      },
      {
        "code": "TVL",
        "name": "Northern Provinces",
        "continent": "AFRICA",
        "region": "Southern Africa"
      },
      {
        "code": "TUN",
        "name": "Tunisia",
        "continent": "AFRICA",
        "region": "Northern Africa"
      },
      {
        "code": "ZAM",
        "name": "Zambia",
        "continent": "AFRICA",
        "region": "South Tropical Africa"
      },
      {
        "code": "ZIM",
        "name": "Zimbabwe",
        "continent": "AFRICA",
        "region": "South Tropical Africa"
      },
      {
        "code": "IRN",
        "name": "Iran",
        "continent": "ASIA-TEMPERATE",
        "region": "Western Asia"
      },
      {
        "code": "LBS",
        "name": "Lebanon-Syria",
        "continent": "ASIA-TEMPERATE",
        "region": "Western Asia"
      },
      {
        "code": "NCS",
        "name": "North Caucasus",
        "continent": "ASIA-TEMPERATE",
        "region": "Caucasus"
      },
      {
        "code": "TCS",
        "name": "Transcaucasus",
        "continent": "ASIA-TEMPERATE",
        "region": "Caucasus"
      },
      {
        "code": "TUR",
        "name": "Türkiye",
        "continent": "ASIA-TEMPERATE",
        "region": "Western Asia"
      },
      {
        "code": "ALB",
        "name": "Albania",
        "continent": "EUROPE",
        "region": "Southeastern Europe"
      },
      {
        "code": "AUT",
        "name": "Austria",
        "continent": "EUROPE",
        "region": "Middle Europe"
      },
      {
        "code": "BGM",
        "name": "Belgium",
        "continent": "EUROPE",
        "region": "Middle Europe"
      },
      {
        "code": "BUL",
        "name": "Bulgaria",
        "continent": "EUROPE",
        "region": "Southeastern Europe"
      },
      {
        "code": "COR",
        "name": "Corse",
        "continent": "EUROPE",
        "region": "Southwestern Europe"
      },
      {
        "code": "CZE",
        "name": "Czechia-Slovakia",
        "continent": "EUROPE",
        "region": "Middle Europe"
      },
      {
        "code": "FRA",
        "name": "France",
        "continent": "EUROPE",
        "region": "Southwestern Europe"
      },
      {
        "code": "GER",
        "name": "Germany",
        "continent": "EUROPE",
        "region": "Middle Europe"
      },
      {
        "code": "GRB",
        "name": "Great Britain",
        "continent": "EUROPE",
        "region": "Northern Europe"
      },
      {
        "code": "GRC",
        "name": "Greece",
        "continent": "EUROPE",
        "region": "Southeastern Europe"
      },
      {
        "code": "HUN",
        "name": "Hungary",
        "continent": "EUROPE",
        "region": "Middle Europe"
      },
      {
        "code": "ITA",
        "name": "Italy",
        "continent": "EUROPE",
        "region": "Southeastern Europe"
      },
      {
        "code": "YUG",
        "name": "NW. Balkan Pen.",
        "continent": "EUROPE",
        "region": "Southeastern Europe"
      },
      {
        "code": "NET",
        "name": "Netherlands",
        "continent": "EUROPE",
        "region": "Middle Europe"
      },
      {
        "code": "POL",
        "name": "Poland",
        "continent": "EUROPE",
        "region": "Middle Europe"
      },
      {
        "code": "POR",
        "name": "Portugal",
        "continent": "EUROPE",
        "region": "Southwestern Europe"
      },
      {
        "code": "ROM",
        "name": "Romania",
        "continent": "EUROPE",
        "region": "Southeastern Europe"
      },
      {
        "code": "SAR",
        "name": "Sardegna",
        "continent": "EUROPE",
        "region": "Southwestern Europe"
      },
      {
        "code": "SPA",
        "name": "Spain",
        "continent": "EUROPE",
        "region": "Southwestern Europe"
      },
      {
        "code": "SWI",
        "name": "Switzerland",
        "continent": "EUROPE",
        "region": "Middle Europe"
      },
      {
        "code": "TUE",
        "name": "Türkiye-in-Europe",
        "continent": "EUROPE",
        "region": "Southeastern Europe"
      },
      {
        "code": "UKR",
        "name": "Ukraine",
        "continent": "EUROPE",
        "region": "Eastern Europe"
      },
      {
        "code": "ALA",
        "name": "Alabama",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "ARI",
        "name": "Arizona",
        "continent": "NORTHERN AMERICA",
        "region": "Southwestern U.S.A."
      },
      {
        "code": "ARK",
        "name": "Arkansas",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "BRC",
        "name": "British Columbia",
        "continent": "NORTHERN AMERICA",
        "region": "Western Canada"
      },
      {
        "code": "CAL",
        "name": "California",
        "continent": "NORTHERN AMERICA",
        "region": "Southwestern U.S.A."
      },
      {
        "code": "CNT",
        "name": "Connecticut",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "DEL",
        "name": "Delaware",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "WDC",
        "name": "District of Columbia",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "FLA",
        "name": "Florida",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "GEO",
        "name": "Georgia",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "IDA",
        "name": "Idaho",
        "continent": "NORTHERN AMERICA",
        "region": "Northwestern U.S.A."
      },
      {
        "code": "ILL",
        "name": "Illinois",
        "continent": "NORTHERN AMERICA",
        "region": "North-Central U.S.A."
      },
      {
        "code": "INI",
        "name": "Indiana",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "IOW",
        "name": "Iowa",
        "continent": "NORTHERN AMERICA",
        "region": "North-Central U.S.A."
      },
      {
        "code": "KAN",
        "name": "Kansas",
        "continent": "NORTHERN AMERICA",
        "region": "North-Central U.S.A."
      },
      {
        "code": "KTY",
        "name": "Kentucky",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "LOU",
        "name": "Louisiana",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "MAI",
        "name": "Maine",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "MRY",
        "name": "Maryland",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "MAS",
        "name": "Massachusetts",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "MXC",
        "name": "Mexico Central",
        "continent": "NORTHERN AMERICA",
        "region": "Mexico"
      },
      {
        "code": "MXG",
        "name": "Mexico Gulf",
        "continent": "NORTHERN AMERICA",
        "region": "Mexico"
      },
      {
        "code": "MXE",
        "name": "Mexico Northeast",
        "continent": "NORTHERN AMERICA",
        "region": "Mexico"
      },
      {
        "code": "MXN",
        "name": "Mexico Northwest",
        "continent": "NORTHERN AMERICA",
        "region": "Mexico"
      },
      {
        "code": "MXT",
        "name": "Mexico Southeast",
        "continent": "NORTHERN AMERICA",
        "region": "Mexico"
      },
      {
        "code": "MXS",
        "name": "Mexico Southwest",
        "continent": "NORTHERN AMERICA",
        "region": "Mexico"
      },
      {
        "code": "MIC",
        "name": "Michigan",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "MIN",
        "name": "Minnesota",
        "continent": "NORTHERN AMERICA",
        "region": "North-Central U.S.A."
      },
      {
        "code": "MSI",
        "name": "Mississippi",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "MSO",
        "name": "Missouri",
        "continent": "NORTHERN AMERICA",
        "region": "North-Central U.S.A."
      },
      {
        "code": "NEB",
        "name": "Nebraska",
        "continent": "NORTHERN AMERICA",
        "region": "North-Central U.S.A."
      },
      {
        "code": "NBR",
        "name": "New Brunswick",
        "continent": "NORTHERN AMERICA",
        "region": "Eastern Canada"
      },
      {
        "code": "NWH",
        "name": "New Hampshire",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "NWJ",
        "name": "New Jersey",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "NWM",
        "name": "New Mexico",
        "continent": "NORTHERN AMERICA",
        "region": "South-Central U.S.A."
      },
      {
        "code": "NWY",
        "name": "New York",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "NFL",
        "name": "Newfoundland",
        "continent": "NORTHERN AMERICA",
        "region": "Eastern Canada"
      },
      {
        "code": "NCA",
        "name": "North Carolina",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "NSC",
        "name": "Nova Scotia",
        "continent": "NORTHERN AMERICA",
        "region": "Eastern Canada"
      },
      {
        "code": "OHI",
        "name": "Ohio",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "OKL",
        "name": "Oklahoma",
        "continent": "NORTHERN AMERICA",
        "region": "North-Central U.S.A."
      },
      {
        "code": "ONT",
        "name": "Ontario",
        "continent": "NORTHERN AMERICA",
        "region": "Eastern Canada"
      },
      {
        "code": "ORE",
        "name": "Oregon",
        "continent": "NORTHERN AMERICA",
        "region": "Northwestern U.S.A."
      },
      {
        "code": "PEN",
        "name": "Pennsylvania",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "QUE",
        "name": "Québec",
        "continent": "NORTHERN AMERICA",
        "region": "Eastern Canada"
      },
      {
        "code": "RHO",
        "name": "Rhode I.",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "SCA",
        "name": "South Carolina",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "TEN",
        "name": "Tennessee",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "TEX",
        "name": "Texas",
        "continent": "NORTHERN AMERICA",
        "region": "South-Central U.S.A."
      },
      {
        "code": "VER",
        "name": "Vermont",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "VRG",
        "name": "Virginia",
        "continent": "NORTHERN AMERICA",
        "region": "Southeastern U.S.A."
      },
      {
        "code": "WAS",
        "name": "Washington",
        "continent": "NORTHERN AMERICA",
        "region": "Northwestern U.S.A."
      },
      {
        "code": "WVA",
        "name": "West Virginia",
        "continent": "NORTHERN AMERICA",
        "region": "Northeastern U.S.A."
      },
      {
        "code": "WIS",
        "name": "Wisconsin",
        "continent": "NORTHERN AMERICA",
        "region": "North-Central U.S.A."
      },
      {
        "code": "CLM",
        "name": "Colombia",
        "continent": "SOUTHERN AMERICA",
        "region": "Western South America"
      },
      {
        "code": "COS",
        "name": "Costa Rica",
        "continent": "SOUTHERN AMERICA",
        "region": "Central America"
      },
      {
        "code": "CUB",
        "name": "Cuba",
        "continent": "SOUTHERN AMERICA",
        "region": "Caribbean"
      },
      {
        "code": "DOM",
        "name": "Dominican Republic",
        "continent": "SOUTHERN AMERICA",
        "region": "Caribbean"
      },
      {
        "code": "GUA",
        "name": "Guatemala",
        "continent": "SOUTHERN AMERICA",
        "region": "Central America"
      },
      {
        "code": "HAI",
        "name": "Haiti",
        "continent": "SOUTHERN AMERICA",
        "region": "Caribbean"
      },
      {
        "code": "JAM",
        "name": "Jamaica",
        "continent": "SOUTHERN AMERICA",
        "region": "Caribbean"
      },
      {
        "code": "PUE",
        "name": "Puerto Rico",
        "continent": "SOUTHERN AMERICA",
        "region": "Caribbean"
      },
      {
        "code": "VEN",
        "name": "Venezuela",
        "continent": "SOUTHERN AMERICA",
        "region": "Northern South America"
      }
    ]
  },
  "phenology": {
    "n": 93,
    "n_annotated": 270,
    "sufficient": true,
    "by_month": [
      9,
      7,
      2,
      3,
      4,
      10,
      16,
      20,
      19,
      1,
      0,
      2
    ],
    "annotated_by_month": [
      17,
      11,
      14,
      13,
      22,
      33,
      32,
      42,
      47,
      28,
      8,
      3
    ],
    "share": [
      0.5294,
      0.6364,
      0.1429,
      0.2308,
      0.1818,
      0.303,
      0.5,
      0.4762,
      0.4043,
      0.0357,
      0,
      null
    ],
    "ci": [
      [
        0.3096,
        0.7384
      ],
      [
        0.3538,
        0.8483
      ],
      [
        0.0401,
        0.3994
      ],
      [
        0.0818,
        0.5026
      ],
      [
        0.0731,
        0.3852
      ],
      [
        0.1738,
        0.4734
      ],
      [
        0.3363,
        0.6637
      ],
      [
        0.3336,
        0.6228
      ],
      [
        0.2764,
        0.5466
      ],
      [
        0.0063,
        0.1771
      ],
      [
        0,
        0.3244
      ],
      null
    ],
    "peak_month": 2,
    "peak_basis": "share",
    "min_month_n": 5,
    "fruiting": {
      "n": 58,
      "by_month": [
        3,
        3,
        1,
        3,
        3,
        7,
        2,
        15,
        12,
        7,
        1,
        1
      ],
      "share": [
        0.1765,
        0.2727,
        0.0714,
        0.2308,
        0.1364,
        0.2121,
        0.0625,
        0.3571,
        0.2553,
        0.25,
        0.125,
        null
      ],
      "ci": [
        [
          0.0619,
          0.4103
        ],
        [
          0.0975,
          0.5657
        ],
        [
          0.0127,
          0.3147
        ],
        [
          0.0818,
          0.5026
        ],
        [
          0.0475,
          0.3334
        ],
        [
          0.1068,
          0.3775
        ],
        [
          0.0173,
          0.2015
        ],
        [
          0.2299,
          0.5083
        ],
        [
          0.1525,
          0.3951
        ],
        [
          0.1268,
          0.4336
        ],
        [
          0.0224,
          0.4709
        ],
        null
      ],
      "peak_month": 8
    },
    "doi": "10.15468/dl.cgje2x",
    "source": "GBIF SQL download of iNaturalist Research-grade Observations",
    "quality_grade": "research (guaranteed upstream by iNaturalist at export)",
    "fields": {
      "by_month": "observations found IN FLOWER, by month (Jan..Dec)",
      "annotated_by_month": "observations in which a human recorded the reproductive state at all, by month. This is the denominator, and it includes the records that said \"no flowers or fruits\".",
      "share": "by_month / annotated_by_month. null where too few plants were examined to state a proportion.",
      "ci": "95% Wilson score interval on share.",
      "peak_month": "the month with the highest SHARE, not the highest count."
    }
  },
  "traits": {
    "lifeform": "subshrub",
    "climate": "temperate"
  },
  "climate_envelope": {
    "mean_annual_temp_c": {
      "p5": 6.4,
      "median": 11.2,
      "p95": 20
    },
    "temp_seasonality": {
      "p5": 281.5,
      "median": 844.2,
      "p95": 1024.2
    },
    "max_temp_warmest_month_c": {
      "p5": 20.8,
      "median": 27.2,
      "p95": 32.8
    },
    "min_temp_coldest_month_c": {
      "p5": -13.6,
      "median": -3.7,
      "p95": 10
    },
    "annual_precip_mm": {
      "p5": 840.7,
      "median": 1182.8,
      "p95": 1718.4
    },
    "precip_seasonality": {
      "p5": 8.7,
      "median": 16.3,
      "p95": 53.4
    },
    "precip_driest_quarter_mm": {
      "p5": 98.8,
      "median": 235.8,
      "p95": 333
    },
    "n": 1978,
    "units": {
      "mean_annual_temp_c": "degrees Celsius",
      "min_temp_coldest_month_c": "degrees Celsius (mean daily MINIMUM of the coldest month)",
      "max_temp_warmest_month_c": "degrees Celsius (mean daily MAXIMUM of the warmest month)",
      "annual_precip_mm": "millimetres per year",
      "precip_driest_quarter_mm": "millimetres in the driest quarter"
    },
    "source": "CHELSA V2.1 climatologies 1981-2010, 30 arcsec",
    "citation": "Karger, D.N. et al. (2017) Climatologies at high resolution for the earth's land surface areas. Scientific Data 4, 170122.",
    "occurrences": "iNaturalist Research-grade Observations",
    "occurrences_doi": "10.15468/dl.cgje2x",
    "method": "Climate sampled at every georeferenced research-grade observation of the species. Reported as the 5th to 95th percentile across 1978 of them. Species with fewer than 30 georeferenced observations get no envelope.",
    "is_hardiness_zone": false,
    "hardiness_zone_note": "min_temp_coldest_month_c is NOT a USDA hardiness zone. A USDA zone is the average annual EXTREME minimum temperature; this is the mean daily minimum of the coldest month, which is typically about 16 C warmer. Do not present one as the other."
  },
  "identification": {
    "on_device_model": {
      "model": "Google AIY plants V1 (TFLite, 2,102 classes)",
      "identifies_this_species": false
    },
    "in_accuracy_benchmark": false,
    "benchmark_url": "https://botanai.app/accuracy/",
    "confused_with": []
  },
  "sources": [
    {
      "name": "World Flora Online Plant List",
      "note": "accepted name, authority, classification",
      "license": "CC0",
      "url": "https://www.worldfloraonline.org/",
      "retrieved": "2026-07-12"
    },
    {
      "name": "iNaturalist",
      "note": "photographs and flowering annotations, CC0 / CC BY / CC BY-SA only",
      "license": "per photograph",
      "url": "https://www.inaturalist.org/",
      "retrieved": "2026-06-27"
    },
    {
      "name": "USDA PLANTS Database",
      "note": "common name, checklist symbol LUPA",
      "license": "public domain",
      "url": "https://plants.usda.gov/",
      "retrieved": "2026-07-13"
    },
    {
      "name": "Kew, World Checklist of Vascular Plants (WCVP v16)",
      "note": "native distribution by TDWG level-3 botanical country, and life form",
      "license": "CC BY 3.0",
      "url": "https://doi.org/10.15468/6h8ucr",
      "retrieved": "2026-06-04"
    }
  ],
  "gaps": [
    {
      "field": "care",
      "status": "not_published",
      "reason": "Not published anywhere on this site. Every care claim we could make cheaply would be an unsourced enum, which is what the previous version of this site did and why it was deleted."
    },
    {
      "field": "toxicity",
      "status": "not_published",
      "reason": "Not published. The widely copied list is a copyrighted compilation restricted to noncommercial use, and we will not scrape it. A sourced toxicity dataset is being built from primary literature and is not ready."
    },
    {
      "field": "botanical_review",
      "status": "not_sourced",
      "reason": "No page on this site has been reviewed by a named botanist. The determination label says so, on every page."
    },
    {
      "field": "hardiness_zone",
      "status": "not_published",
      "reason": "Not published for any species. A USDA hardiness zone is the average annual EXTREME minimum temperature, and we hold the mean daily minimum of the coldest month, which is a different quantity and typically about 16 C warmer. Presenting one as the other would place a plant several zones too warm, so we do not publish a zone at all."
    }
  ],
  "license": {
    "data": "CC BY 4.0",
    "attribution": "BotanAI, https://botanai.app/plants/ludwigia-palustris/",
    "note": "This document is licensed CC BY 4.0. The PHOTOGRAPHS it links are licensed individually and separately: each carries its own license and license_url, and those terms govern. Underlying taxonomy is CC0 (World Flora Online); native range is CC BY 3.0 (Kew)."
  }
}