{
  "$schema": "https://botanai.app/schema/species-v1.json",
  "schema_version": 1,
  "url": "https://botanai.app/plants/lopholejeunea-nigricans/",
  "last_modified": "2026-06-03",
  "id": {
    "slug": "lopholejeunea-nigricans",
    "wfo": "wfo-0001208251",
    "inat_taxon_id": 327495
  },
  "name": {
    "scientific": "Lopholejeunea nigricans",
    "authorship": "(Lindenb.) Steph. ex Schiffn.",
    "genus": "Lopholejeunea",
    "epithet": "nigricans",
    "family": "Lejeuneaceae",
    "rank": "species",
    "taxonomic_status": "accepted",
    "accepted_according_to": "World Flora Online Plant List, 2026-06",
    "is_homonym": false,
    "published_in": "Consp. Hepat. Arch. Ind.: 293. 1898.",
    "protologue_url": "https://www.bryonames.org/",
    "common": {
      "preferred": null,
      "english": [],
      "source": null
    },
    "synonyms": [
      {
        "name": "Acrolejeunea atroviridis",
        "authorship": "(Spruce) Steph."
      },
      {
        "name": "Archilejeunea palustris",
        "authorship": "Sim"
      },
      {
        "name": "Brachiolejeunea gibbosa",
        "authorship": "(Ångstr.) Steph."
      },
      {
        "name": "Heterolejeunea javanica",
        "authorship": "Schiffn."
      },
      {
        "name": "Heterolejeunea javanica f. aquatica",
        "authorship": "Schiffn."
      },
      {
        "name": "Lejeunea abortiva",
        "authorship": "Mitt."
      },
      {
        "name": "Lejeunea atroviridis",
        "authorship": "Spruce"
      },
      {
        "name": "Lejeunea gibbosa",
        "authorship": "Ångstr."
      },
      {
        "name": "Lejeunea intermedia",
        "authorship": "Lindenb."
      },
      {
        "name": "Lejeunea javanica",
        "authorship": "Nees"
      },
      {
        "name": "Lejeunea mannii",
        "authorship": "Austin"
      },
      {
        "name": "Lejeunea mariei",
        "authorship": "Besch. & Spruce"
      },
      {
        "name": "Lejeunea muelleriana",
        "authorship": "Gottsche"
      },
      {
        "name": "Lejeunea multilacera",
        "authorship": "(Steph.) J.M.Coult., Barnes & Arthur"
      },
      {
        "name": "Lejeunea nigricans",
        "authorship": "Lindenb."
      },
      {
        "name": "Lejeunea rusbyi",
        "authorship": "Spruce"
      },
      {
        "name": "Lejeunea subinermis",
        "authorship": "(Steph.) Steph."
      },
      {
        "name": "Lopholejeunea abortiva",
        "authorship": "(Mitt.) Steph."
      },
      {
        "name": "Lopholejeunea abortiva var. alata",
        "authorship": "Vanden Berghen"
      },
      {
        "name": "Lopholejeunea abortiva var. doliiformis",
        "authorship": "U.S.Awasthi, S.C.Srivast. & D.Sharma"
      },
      {
        "name": "Lopholejeunea abortiva var. fragilis",
        "authorship": "(Steph.) Vanden Berghen"
      },
      {
        "name": "Lopholejeunea acutifolia",
        "authorship": "Mizut. & Piippo"
      },
      {
        "name": "Lopholejeunea aemula",
        "authorship": "Schiffn."
      },
      {
        "name": "Lopholejeunea andersonii",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea apiahyna",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea apiculata",
        "authorship": "Horik."
      },
      {
        "name": "Lopholejeunea atroviridis",
        "authorship": "(Spruce) A.Evans"
      },
      {
        "name": "Lopholejeunea australis",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea brunnea",
        "authorship": "Horik."
      },
      {
        "name": "Lopholejeunea fragilis",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea gibbosa",
        "authorship": "(Ångstr.) Steph."
      },
      {
        "name": "Lopholejeunea gradsteinii",
        "authorship": "Udar, U.S.Awasthi & Shaheen"
      },
      {
        "name": "Lopholejeunea hawaica",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea intermedia",
        "authorship": "(Lindenb.) Steph."
      },
      {
        "name": "Lopholejeunea javanica",
        "authorship": "(Nees) Schiffn."
      },
      {
        "name": "Lopholejeunea kashyapii",
        "authorship": "U.S.Awasthi, S.C.Srivast. & D.Sharma"
      },
      {
        "name": "Lopholejeunea kilimanjarica",
        "authorship": "E.W.Jones"
      },
      {
        "name": "Lopholejeunea levieriana",
        "authorship": "C.Massal."
      },
      {
        "name": "Lopholejeunea mannii",
        "authorship": "(Austin) Steph."
      },
      {
        "name": "Lopholejeunea mariei",
        "authorship": "(Besch. & Spruce) Steph."
      },
      {
        "name": "Lopholejeunea mitis",
        "authorship": "Herzog"
      },
      {
        "name": "Lopholejeunea muelleriana",
        "authorship": "(Gottsche) Schiffn."
      },
      {
        "name": "Lopholejeunea muelleriana subsp. floridana",
        "authorship": "R.M.Schust."
      },
      {
        "name": "Lopholejeunea muelleriana subsp. puertoricensis",
        "authorship": "R.M.Schust."
      },
      {
        "name": "Lopholejeunea muelleriana var. australis",
        "authorship": "(Steph.) B.M.Thiers & Gradst."
      },
      {
        "name": "Lopholejeunea muensis",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea muensis var. microloba",
        "authorship": "Pearson"
      },
      {
        "name": "Lopholejeunea multiflora",
        "authorship": "J.B.Jack & Steph."
      },
      {
        "name": "Lopholejeunea multilacera",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea nigricans var. intermedia",
        "authorship": "(Lindenb.) Schiffn."
      },
      {
        "name": "Lopholejeunea nigricans var. nigricans",
        "authorship": null
      },
      {
        "name": "Lopholejeunea owahuensis",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea palustris",
        "authorship": "(Sim) S.W.Arnell"
      },
      {
        "name": "Lopholejeunea parva",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea parvistipula",
        "authorship": "Herzog"
      },
      {
        "name": "Lopholejeunea proxima",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea quinquecarinata",
        "authorship": "Vanden Berghen"
      },
      {
        "name": "Lopholejeunea rivularis",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea rusbyi",
        "authorship": "(Spruce) Steph."
      },
      {
        "name": "Lopholejeunea spinosa",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea subinermis",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea subnuda",
        "authorship": "(Mitt.) Steph."
      },
      {
        "name": "Lopholejeunea tonkinensis",
        "authorship": "Steph."
      },
      {
        "name": "Lopholejeunea vermoesenii",
        "authorship": "Pearson"
      },
      {
        "name": "Lopholejeunea zenkeri",
        "authorship": "Steph."
      },
      {
        "name": "Mastigolejeunea honoluluana",
        "authorship": "Steph."
      },
      {
        "name": "Neurolejeunea lechleri",
        "authorship": "Steph."
      },
      {
        "name": "Phragmicoma subnuda",
        "authorship": "Mitt."
      },
      {
        "name": "Ptychocoleus atroviridis",
        "authorship": "(Spruce) Steph."
      },
      {
        "name": "Symbiezidium intermedium",
        "authorship": "(Lindenb.) Trevis."
      },
      {
        "name": "Symbiezidium javanicum",
        "authorship": "(Nees) Trevis."
      },
      {
        "name": "Symbiezidium nigricans",
        "authorship": "(Lindenb.) Trevis."
      }
    ],
    "wikipedia": null
  },
  "photographs": {
    "shown": 3,
    "usable_found": 3,
    "licences": [
      "CC-BY"
    ],
    "noncommercial_excluded": true,
    "items": [
      {
        "photo_id": 188653830,
        "license": "CC-BY",
        "license_name": "CC BY 4.0",
        "license_url": "https://creativecommons.org/licenses/by/4.0/",
        "derivatives_restricted": false,
        "creator": "Kevin Faccenda",
        "creator_url": "https://www.inaturalist.org/people/kevinfaccenda",
        "source_url": "https://www.inaturalist.org/observations/d50c3f51-f118-46e5-a173-c1a555b43656",
        "observed_on": "2022-04-02",
        "lat": 21.338,
        "lon": -157.811,
        "original": {
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          "jpeg": {
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            "800": "https://img.botanai.app/p/188653830/r1/800.jpg"
          }
        },
        "attribution": "Kevin Faccenda, CC BY 4.0, via iNaturalist"
      },
      {
        "photo_id": 180747907,
        "license": "CC-BY",
        "license_name": "CC BY 4.0",
        "license_url": "https://creativecommons.org/licenses/by/4.0/",
        "derivatives_restricted": false,
        "creator": "Kevin Faccenda",
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        "source_url": "https://www.inaturalist.org/observations/1aaf59c4-fe1f-471a-a8a8-88a70bbcd2c2",
        "observed_on": "2022-02-20",
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          "jpeg": {
            "400": "https://img.botanai.app/p/180747907/r1/400.jpg",
            "800": "https://img.botanai.app/p/180747907/r1/800.jpg"
          }
        },
        "attribution": "Kevin Faccenda, CC BY 4.0, via iNaturalist"
      },
      {
        "photo_id": 180747899,
        "license": "CC-BY",
        "license_name": "CC BY 4.0",
        "license_url": "https://creativecommons.org/licenses/by/4.0/",
        "derivatives_restricted": false,
        "creator": "Kevin Faccenda",
        "creator_url": "https://www.inaturalist.org/people/kevinfaccenda",
        "source_url": "https://www.inaturalist.org/observations/1aaf59c4-fe1f-471a-a8a8-88a70bbcd2c2",
        "observed_on": "2022-02-20",
        "lat": 21.358,
        "lon": -157.815,
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        },
        "widths": [
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        ],
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          "jpeg": {
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        "attribution": "Kevin Faccenda, CC BY 4.0, via iNaturalist"
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    ]
  },
  "native_range": null,
  "phenology": null,
  "traits": {
    "lifeform": null,
    "climate": null
  },
  "climate_envelope": {
    "mean_annual_temp_c": {
      "p5": 17.5,
      "median": 21.3,
      "p95": 22.8
    },
    "temp_seasonality": {
      "p5": 99.5,
      "median": 120.5,
      "p95": 136.6
    },
    "max_temp_warmest_month_c": {
      "p5": 21.1,
      "median": 23.9,
      "p95": 26.3
    },
    "min_temp_coldest_month_c": {
      "p5": 13.4,
      "median": 18.6,
      "p95": 19.9
    },
    "annual_precip_mm": {
      "p5": 1057.8,
      "median": 1194.7,
      "p95": 4100.7
    },
    "precip_seasonality": {
      "p5": 12,
      "median": 24.2,
      "p95": 49.1
    },
    "precip_driest_quarter_mm": {
      "p5": 135.2,
      "median": 248.5,
      "p95": 786.7
    },
    "n": 40,
    "units": {
      "mean_annual_temp_c": "degrees Celsius",
      "min_temp_coldest_month_c": "degrees Celsius (mean daily MINIMUM of the coldest month)",
      "max_temp_warmest_month_c": "degrees Celsius (mean daily MAXIMUM of the warmest month)",
      "annual_precip_mm": "millimetres per year",
      "precip_driest_quarter_mm": "millimetres in the driest quarter"
    },
    "source": "CHELSA V2.1 climatologies 1981-2010, 30 arcsec",
    "citation": "Karger, D.N. et al. (2017) Climatologies at high resolution for the earth's land surface areas. Scientific Data 4, 170122.",
    "occurrences": "iNaturalist Research-grade Observations",
    "occurrences_doi": null,
    "method": "Climate sampled at every georeferenced research-grade observation of the species. Reported as the 5th to 95th percentile across 40 of them. Species with fewer than 30 georeferenced observations get no envelope.",
    "is_hardiness_zone": false,
    "hardiness_zone_note": "min_temp_coldest_month_c is NOT a USDA hardiness zone. A USDA zone is the average annual EXTREME minimum temperature; this is the mean daily minimum of the coldest month, which is typically about 16 C warmer. Do not present one as the other."
  },
  "identification": {
    "on_device_model": {
      "model": "Google AIY plants V1 (TFLite, 2,102 classes)",
      "identifies_this_species": false
    },
    "in_accuracy_benchmark": false,
    "benchmark_url": "https://botanai.app/accuracy/",
    "confused_with": []
  },
  "sources": [
    {
      "name": "World Flora Online Plant List",
      "note": "accepted name, authority, classification",
      "license": "CC0",
      "url": "https://www.worldfloraonline.org/",
      "retrieved": "2026-07-12"
    },
    {
      "name": "iNaturalist",
      "note": "photographs and flowering annotations, CC0 / CC BY / CC BY-SA only",
      "license": "per photograph",
      "url": "https://www.inaturalist.org/",
      "retrieved": "2026-06-27"
    }
  ],
  "gaps": [
    {
      "field": "care",
      "status": "not_published",
      "reason": "Not published anywhere on this site. Every care claim we could make cheaply would be an unsourced enum, which is what the previous version of this site did and why it was deleted."
    },
    {
      "field": "toxicity",
      "status": "not_published",
      "reason": "Not published. The widely copied list is a copyrighted compilation restricted to noncommercial use, and we will not scrape it. A sourced toxicity dataset is being built from primary literature and is not ready."
    },
    {
      "field": "botanical_review",
      "status": "not_sourced",
      "reason": "No page on this site has been reviewed by a named botanist. The determination label says so, on every page."
    },
    {
      "field": "native_range",
      "status": "not_sourced",
      "reason": "Kew's World Checklist of Vascular Plants does not cover this taxon."
    },
    {
      "field": "common_name",
      "status": "not_sourced",
      "reason": "No vernacular name for this taxon in Wikidata (P1843) or iNaturalist. We do not invent one, and we do not present the scientific name as though it were a common name."
    },
    {
      "field": "hardiness_zone",
      "status": "not_published",
      "reason": "Not published for any species. A USDA hardiness zone is the average annual EXTREME minimum temperature, and we hold the mean daily minimum of the coldest month, which is a different quantity and typically about 16 C warmer. Presenting one as the other would place a plant several zones too warm, so we do not publish a zone at all."
    },
    {
      "field": "phenology",
      "status": "not_sourced",
      "reason": "No dated flowering annotations found for this taxon."
    }
  ],
  "license": {
    "data": "CC BY 4.0",
    "attribution": "BotanAI, https://botanai.app/plants/lopholejeunea-nigricans/",
    "note": "This document is licensed CC BY 4.0. The PHOTOGRAPHS it links are licensed individually and separately: each carries its own license and license_url, and those terms govern. Underlying taxonomy is CC0 (World Flora Online); native range is CC BY 3.0 (Kew)."
  }
}