Leptogramma tottoidesH.Itô

WFO wfo-0001116094 Accepted WFO 2026-06 8 photographs CC BY

Plate 1 figs. a–h · 4 observations

This species has been photographed under an open licence only 4 times, so some figures below are different views of the same plant, taken on the same day, rather than different individuals. They are usually different parts of it: the leaf, the flower, the bark.

Leptogramma tottoides, photographed by Jacy Chen
fig. a Jacy Chen, CC BY 4.0 / 2021-10-31 / obs. 169179773
Leptogramma tottoides, photographed by Jacy Chen
fig. b Jacy Chen, CC BY 4.0 / obs. 163436486
Leptogramma tottoides, photographed by Jacy Chen
fig. c Jacy Chen, CC BY 4.0 / obs. 135391430
Leptogramma tottoides, photographed by Jacy Chen
fig. d Jacy Chen, CC BY 4.0 / obs. 105589764
Leptogramma tottoides, photographed by Jacy Chen
fig. e Jacy Chen, CC BY 4.0 / obs. 169179760
Leptogramma tottoides, photographed by Jacy Chen
fig. f Jacy Chen, CC BY 4.0 / obs. 135391359
Leptogramma tottoides, photographed by Jacy Chen
fig. g Jacy Chen, CC BY 4.0 / obs. 135391382
Leptogramma tottoides, photographed by Jacy Chen
fig. h Jacy Chen, CC BY 4.0 / obs. 163436516

Every figure is a research-grade observation under CC0, CC BY or CC BY-SA, rehosted with the photographer’s name, the licence and the observation it came from. Photographs under a NonCommercial licence are excluded from this site and are never stored, which costs us a great many pictures and is not negotiable.

Native range 3 botanical countries

Regions where Leptogramma tottoides is native: China South-Central, China Southeast, Taiwan China South-CentralChina SoutheastTaiwan
Native distribution of Leptogramma tottoides, after Kew’s World Checklist of Vascular Plants. Introduced, extinct and doubtful records are excluded, so this is where the plant is from, not everywhere it now grows.
RegionTDWG codeContinent
China South-Central CHC ASIA-TEMPERATE
China Southeast CHS
Taiwan TAI

Region boundaries approximated from Natural Earth (public domain) and mapped to TDWG World Geographical Scheme for Recording Plant Distributions (WGSRPD) level-3 botanical countries (Brummitt 2001). Indicative, not the official WGSRPD geometry.

Where it actually grows measured, from 52 observations

Condition 5th percentile Median 95th percentile
Coldest month, mean daily low -0.4 °C 3.1 °C 10.1 °C
Warmest month, mean daily high 16.8 °C 21.0 °C 26.8 °C
Annual rainfall 2,981 mm 4,144 mm 4,790 mm
Rainfall in the driest quarter 183 mm 289 mm 759 mm

It is found where winters bring light frost. This is not care advice and it is not a forecast. It is a measurement: we sampled the climate at every one of the 52 research-grade observations of Leptogramma tottoides that carry a coordinate, and this is the range those places actually span. The 5th and 95th percentiles are used rather than the minimum and maximum, because a single cultivated specimen in a heated conservatory should not widen a tropical plant's range to the Arctic.

This is not a hardiness zone. A USDA zone is the average annual extreme minimum temperature. The figure above is the mean daily minimum of the coldest month, which is a different quantity and is typically far warmer. Reading one as the other would place a plant several zones too warm, so we do not publish a hardiness zone, because we do not have one.

Also published as 7 synonyms

A synonym is not an error. It is a record of botanists disagreeing, in print, about where this plant belongs. Each of these was somebody’s considered answer.

  • Cyclosorus tottoides (H.Itô) C.M.Kuo
  • Leptogramma caudata Ching
  • Leptogramma totta var. tottoides (Hayata ex H.Itô) H.Itô
  • Stegnogramma tottoides (H.Itô) K.Iwats.
  • Thelypteris caudata (Ching) C.F.Reed
  • Thelypteris tottoides (Hayata ex H.Itô) C.V.Morton
  • Thelypteris tottoides (Hayata ex H.Itô) C.M.Kuo

Sourcesevery claim on this page

  1. World Flora Online Plant List. accepted name, authority, classification. CC0. Retrieved 2026-07-12.
  2. iNaturalist. photographs and flowering annotations, CC0 / CC BY / CC BY-SA only. per photograph. Retrieved 2026-06-27.
  3. Kew, World Checklist of Vascular Plants (WCVP v16). native distribution by TDWG level-3 botanical country, and life form. CC BY 3.0. Retrieved 2026-06-04.

We publish what we can source and we say so when we cannot. This page has no care advice and no toxicity claim, because we do not yet have those from a source we can cite.