{
  "$schema": "https://botanai.app/schema/species-v1.json",
  "schema_version": 1,
  "url": "https://botanai.app/plants/geheebia-tophacea/",
  "last_modified": "2026-06-03",
  "id": {
    "slug": "geheebia-tophacea",
    "wfo": "wfo-1000018273",
    "inat_taxon_id": 1282051
  },
  "name": {
    "scientific": "Geheebia tophacea",
    "authorship": "(Brid.) R.H.Zander",
    "genus": "Geheebia",
    "epithet": "tophacea",
    "family": "Pottiaceae",
    "rank": "species",
    "taxonomic_status": "accepted",
    "accepted_according_to": "World Flora Online Plant List, 2026-06",
    "is_homonym": false,
    "published_in": "Framew. Post-Phylogenet. Syst.: 95. 2013.",
    "protologue_url": "https://www.bryonames.org/",
    "common": {
      "preferred": "didymodon moss",
      "english": [
        "didymodon moss"
      ],
      "source": "usda"
    },
    "synonyms": [
      {
        "name": "Anacalypta tophacea",
        "authorship": "(Brid.) Nees & Hornsch."
      },
      {
        "name": "Barbula abbonii",
        "authorship": "Thér."
      },
      {
        "name": "Barbula brevifolia",
        "authorship": "Lindb."
      },
      {
        "name": "Barbula brevifolia var. acutifolia",
        "authorship": "Braithw."
      },
      {
        "name": "Barbula brevifolia var. lingulata",
        "authorship": "(Boulay) C.E.O.Jensen"
      },
      {
        "name": "Barbula excurrens",
        "authorship": "Broth."
      },
      {
        "name": "Barbula pringlei",
        "authorship": "(Cardot) Hilp."
      },
      {
        "name": "Barbula spadicea var. silurica",
        "authorship": "(Velen.) Podp."
      },
      {
        "name": "Barbula tophacea",
        "authorship": "(Brid.) Mitt."
      },
      {
        "name": "Barbula tophacea f. acutifolia",
        "authorship": "Mönk."
      },
      {
        "name": "Barbula tophacea f. anatina",
        "authorship": "(Hamm.) Podp."
      },
      {
        "name": "Barbula tophacea f. bosniaca",
        "authorship": "(Głow.) Mönk."
      },
      {
        "name": "Barbula tophacea f. cylindrica",
        "authorship": "(Boulay) Podp."
      },
      {
        "name": "Barbula tophacea f. elata",
        "authorship": "(Boulay) Mönk."
      },
      {
        "name": "Barbula tophacea f. humilis",
        "authorship": "(Schimp.) Mönk."
      },
      {
        "name": "Barbula tophacea f. laxa",
        "authorship": "(Kindb.) Podp."
      },
      {
        "name": "Barbula tophacea f. linearis",
        "authorship": "(Limpr.) Podp."
      },
      {
        "name": "Barbula tophacea f. lingulata",
        "authorship": "(Boulay) Mönk."
      },
      {
        "name": "Barbula tophacea f. luxurians",
        "authorship": "Dixon"
      },
      {
        "name": "Barbula tophacea f. propagulifera",
        "authorship": "(J.J.Amann) Mönk."
      },
      {
        "name": "Barbula tophacea f. recurvifolia",
        "authorship": "(Boulay) Demaret"
      },
      {
        "name": "Barbula tophacea f. scabrinervis",
        "authorship": "(Podp.) Podp."
      },
      {
        "name": "Barbula tophacea f. thermarum",
        "authorship": "Boros"
      },
      {
        "name": "Barbula tophacea f. torrentium",
        "authorship": "(Loeske) Podp."
      },
      {
        "name": "Barbula tophacea f. truncata",
        "authorship": "(Boulay) Podp."
      },
      {
        "name": "Barbula tophacea var. acutifolia",
        "authorship": "Dixon"
      },
      {
        "name": "Barbula tophacea var. breidleri",
        "authorship": "(E.Bauer) Podp."
      },
      {
        "name": "Barbula tophacea var. brevifolia",
        "authorship": "(Bruch & Schimp.) Dixon"
      },
      {
        "name": "Barbula tophacea var. excurrens",
        "authorship": "(Broth.) Podp."
      },
      {
        "name": "Barbula tophacea var. riparia",
        "authorship": "(J.J.Amann) Podp."
      },
      {
        "name": "Barbula tophacea var. torrentium",
        "authorship": "Loeske"
      },
      {
        "name": "Barbula wildii",
        "authorship": "Broth."
      },
      {
        "name": "Ceratodon purpureus var. mammillosus",
        "authorship": "Warnst."
      },
      {
        "name": "Dactylhymenium pringlei",
        "authorship": "Cardot"
      },
      {
        "name": "Desmatodon hendersonii",
        "authorship": "(Renauld & Cardot) R.S.Williams"
      },
      {
        "name": "Desmatodon lingulatus",
        "authorship": "(Hook.f. & Wilson) Sainsbury"
      },
      {
        "name": "Didymodon bosniacus",
        "authorship": "Głow."
      },
      {
        "name": "Didymodon diversifolius",
        "authorship": "Austin"
      },
      {
        "name": "Didymodon excurrens",
        "authorship": "(Broth.) Broth."
      },
      {
        "name": "Didymodon hendersonii",
        "authorship": "Renauld & Cardot"
      },
      {
        "name": "Didymodon ligulifolius",
        "authorship": "G.Roth"
      },
      {
        "name": "Didymodon lingulatus",
        "authorship": "(Hook.f. & Wilson) Broth."
      },
      {
        "name": "Didymodon planifolius",
        "authorship": "J.Froehl."
      },
      {
        "name": "Didymodon planotophaceus",
        "authorship": "J.Froehl."
      },
      {
        "name": "Didymodon riparius",
        "authorship": "J.J.Amann"
      },
      {
        "name": "Didymodon spadiceus var. siluricus",
        "authorship": "Velen."
      },
      {
        "name": "Didymodon tophaceus",
        "authorship": "(Brid.) Lisa"
      },
      {
        "name": "Didymodon tophaceus f. acutifolius",
        "authorship": "Zodda"
      },
      {
        "name": "Didymodon tophaceus f. anatinus",
        "authorship": "(Hamm.) Margad."
      },
      {
        "name": "Didymodon tophaceus f. brevicaulis",
        "authorship": "Zodda"
      },
      {
        "name": "Didymodon tophaceus f. cylindricus",
        "authorship": "(Boulay) Limpr."
      },
      {
        "name": "Didymodon tophaceus f. elatus",
        "authorship": "(Boulay) Artaria ex E.Bauer"
      },
      {
        "name": "Didymodon tophaceus f. humilis",
        "authorship": "(Schimp.) Margad."
      },
      {
        "name": "Didymodon tophaceus f. laxus",
        "authorship": "(Kindb.) R.H.Zander"
      },
      {
        "name": "Didymodon tophaceus f. ligulifolius",
        "authorship": "(G.Roth) Margad."
      },
      {
        "name": "Didymodon tophaceus f. lingulatus",
        "authorship": "(Boulay) Mönk."
      },
      {
        "name": "Didymodon tophaceus f. propagulifer",
        "authorship": "J.J.Amann"
      },
      {
        "name": "Didymodon tophaceus f. recurvifolius",
        "authorship": "(Boulay) De Wild."
      },
      {
        "name": "Didymodon tophaceus f. scabrinervis",
        "authorship": "Podp."
      },
      {
        "name": "Didymodon tophaceus f. truncatus",
        "authorship": "(Boulay) Limpr."
      },
      {
        "name": "Didymodon tophaceus f. vulgaris",
        "authorship": "Mönk."
      },
      {
        "name": "Didymodon tophaceus subsp. ligulifolius",
        "authorship": "(G.Roth) J.J.Amann"
      },
      {
        "name": "Didymodon tophaceus subsp. tophaceus",
        "authorship": null
      },
      {
        "name": "Didymodon tophaceus var. acutifolius",
        "authorship": "Limpr."
      },
      {
        "name": "Didymodon tophaceus var. anatinus",
        "authorship": "Hamm."
      },
      {
        "name": "Didymodon tophaceus var. bosniacus",
        "authorship": "(Głow.) Latzel"
      },
      {
        "name": "Didymodon tophaceus var. breidleri",
        "authorship": "E.Bauer"
      },
      {
        "name": "Didymodon tophaceus var. brevicaulis",
        "authorship": "(Schimp.) Latzel"
      },
      {
        "name": "Didymodon tophaceus var. brevifolius",
        "authorship": "(Bruch & Schimp.) Limpr."
      },
      {
        "name": "Didymodon tophaceus var. cucullatus",
        "authorship": "Bott."
      },
      {
        "name": "Didymodon tophaceus var. decurrens",
        "authorship": "Cardot & Thér."
      },
      {
        "name": "Didymodon tophaceus var. elatus",
        "authorship": "(Boulay) E.Bauer"
      },
      {
        "name": "Didymodon tophaceus var. excurrens",
        "authorship": "(Broth.) Wijk & Margad."
      },
      {
        "name": "Didymodon tophaceus var. humilis",
        "authorship": "(Schimp.) Limpr."
      },
      {
        "name": "Didymodon tophaceus var. laxus",
        "authorship": "(Kindb.) Paris"
      },
      {
        "name": "Didymodon tophaceus var. ligulifolius",
        "authorship": "(G.Roth) J.J.Amann"
      },
      {
        "name": "Didymodon tophaceus var. linearis",
        "authorship": "Limpr."
      },
      {
        "name": "Didymodon tophaceus var. lingulatus",
        "authorship": "(Boulay) Warnst."
      },
      {
        "name": "Didymodon tophaceus var. riparius",
        "authorship": "J.J.Amann"
      },
      {
        "name": "Didymodon tophaceus var. scabrinervis",
        "authorship": "(Podp.) Pavletić"
      },
      {
        "name": "Didymodon tophaceus var. tophaceus",
        "authorship": null
      },
      {
        "name": "Didymodon trifarius var. tophaceus",
        "authorship": "(Brid.) Mont."
      },
      {
        "name": "Didymodon wildii",
        "authorship": "(Broth.) Broth."
      },
      {
        "name": "Gymnostomum platyphyllum",
        "authorship": "Kindb."
      },
      {
        "name": "Gymnostomum wageri",
        "authorship": "Schelpe"
      },
      {
        "name": "Gyroweisia boliviana",
        "authorship": "R.S.Williams"
      },
      {
        "name": "Husnotiella baueri",
        "authorship": "E.B.Bartram"
      },
      {
        "name": "Husnotiella pringlei",
        "authorship": "(Cardot) Grout"
      },
      {
        "name": "Hymenostylium platyphyllum",
        "authorship": "(Kindb.) Broth."
      },
      {
        "name": "Trichostomum excurrens",
        "authorship": "(Broth.) Kindb."
      },
      {
        "name": "Trichostomum knightii",
        "authorship": "Hampe ex Müll.Hal."
      },
      {
        "name": "Trichostomum limosum",
        "authorship": "Dozy & Molk."
      },
      {
        "name": "Trichostomum lingulatum",
        "authorship": "Hook.f. & Wilson"
      },
      {
        "name": "Trichostomum tophaceum",
        "authorship": "Brid."
      },
      {
        "name": "Trichostomum tophaceum f. acutifolium",
        "authorship": "Boulay"
      },
      {
        "name": "Trichostomum tophaceum f. brevicaule",
        "authorship": "Boulay"
      },
      {
        "name": "Trichostomum tophaceum f. cylindricum",
        "authorship": "Boulay"
      },
      {
        "name": "Trichostomum tophaceum f. elatum",
        "authorship": "Boulay"
      },
      {
        "name": "Trichostomum tophaceum f. lingulatum",
        "authorship": "Boulay"
      },
      {
        "name": "Trichostomum tophaceum f. recurvifolium",
        "authorship": "Boulay"
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      {
        "name": "Trichostomum tophaceum f. truncatum",
        "authorship": "Boulay"
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      {
        "name": "Trichostomum tophaceum var. acutifolium",
        "authorship": "Schimp."
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      {
        "name": "Trichostomum tophaceum var. brevicaule",
        "authorship": "Schimp."
      },
      {
        "name": "Trichostomum tophaceum var. brevifolium",
        "authorship": "Bruch & Schimp."
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      {
        "name": "Trichostomum tophaceum var. humile",
        "authorship": "Schimp."
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      {
        "name": "Trichostomum tophaceum var. laxum",
        "authorship": "Kindb."
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      {
        "name": "Trichostomum tophaceum var. leucotrichion",
        "authorship": "Brid."
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      {
        "name": "Trichostomum trifarium var. sterile",
        "authorship": "Fior.-Mazz."
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      {
        "name": "Trichostomum trifarium var. tophaceum",
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      {
        "name": "Trichostomum wildii",
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      {
        "name": "Weissia amblyphylla",
        "authorship": "J.E.Zetterst."
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      {
        "name": "Weissia platyphylla",
        "authorship": "(Kindb.) Kindb."
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        "creator": "Helen Waterman",
        "creator_url": "https://www.inaturalist.org/people/helenwaterman",
        "source_url": "https://www.inaturalist.org/observations/0a262bbb-2917-4f54-bde3-cfa0819e1031",
        "observed_on": "2022-05-21",
        "lat": 53.383,
        "lon": -1.835,
        "original": {
          "width": 622,
          "height": 1024
        },
        "widths": [
          400
        ],
        "urls": {
          "avif": {
            "400": "https://img.botanai.app/p/199911343/r1/400.avif"
          },
          "jpeg": {
            "400": "https://img.botanai.app/p/199911343/r1/400.jpg"
          }
        },
        "attribution": "Helen Waterman, CC BY 4.0, via iNaturalist"
      }
    ]
  },
  "native_range": null,
  "phenology": null,
  "traits": {
    "lifeform": null,
    "climate": null
  },
  "climate_envelope": {
    "mean_annual_temp_c": {
      "p5": 6.8,
      "median": 10.2,
      "p95": 16
    },
    "temp_seasonality": {
      "p5": 357.5,
      "median": 544.3,
      "p95": 885.4
    },
    "max_temp_warmest_month_c": {
      "p5": 17.5,
      "median": 22.2,
      "p95": 32.3
    },
    "min_temp_coldest_month_c": {
      "p5": -8.5,
      "median": -0.1,
      "p95": 7.4
    },
    "annual_precip_mm": {
      "p5": 344.8,
      "median": 795.9,
      "p95": 1393.6
    },
    "precip_seasonality": {
      "p5": 11.8,
      "median": 24.6,
      "p95": 93
    },
    "precip_driest_quarter_mm": {
      "p5": 6.9,
      "median": 149.9,
      "p95": 251.6
    },
    "n": 33,
    "units": {
      "mean_annual_temp_c": "degrees Celsius",
      "min_temp_coldest_month_c": "degrees Celsius (mean daily MINIMUM of the coldest month)",
      "max_temp_warmest_month_c": "degrees Celsius (mean daily MAXIMUM of the warmest month)",
      "annual_precip_mm": "millimetres per year",
      "precip_driest_quarter_mm": "millimetres in the driest quarter"
    },
    "source": "CHELSA V2.1 climatologies 1981-2010, 30 arcsec",
    "citation": "Karger, D.N. et al. (2017) Climatologies at high resolution for the earth's land surface areas. Scientific Data 4, 170122.",
    "occurrences": "iNaturalist Research-grade Observations",
    "occurrences_doi": null,
    "method": "Climate sampled at every georeferenced research-grade observation of the species. Reported as the 5th to 95th percentile across 33 of them. Species with fewer than 30 georeferenced observations get no envelope.",
    "is_hardiness_zone": false,
    "hardiness_zone_note": "min_temp_coldest_month_c is NOT a USDA hardiness zone. A USDA zone is the average annual EXTREME minimum temperature; this is the mean daily minimum of the coldest month, which is typically about 16 C warmer. Do not present one as the other."
  },
  "identification": {
    "on_device_model": {
      "model": "Google AIY plants V1 (TFLite, 2,102 classes)",
      "identifies_this_species": false
    },
    "in_accuracy_benchmark": false,
    "benchmark_url": "https://botanai.app/accuracy/",
    "confused_with": []
  },
  "sources": [
    {
      "name": "World Flora Online Plant List",
      "note": "accepted name, authority, classification",
      "license": "CC0",
      "url": "https://www.worldfloraonline.org/",
      "retrieved": "2026-07-12"
    },
    {
      "name": "iNaturalist",
      "note": "photographs and flowering annotations, CC0 / CC BY / CC BY-SA only",
      "license": "per photograph",
      "url": "https://www.inaturalist.org/",
      "retrieved": "2026-06-27"
    },
    {
      "name": "USDA PLANTS Database",
      "note": "common name, checklist symbol DITO4",
      "license": "public domain",
      "url": "https://plants.usda.gov/",
      "retrieved": "2026-07-13"
    }
  ],
  "gaps": [
    {
      "field": "care",
      "status": "not_published",
      "reason": "Not published anywhere on this site. Every care claim we could make cheaply would be an unsourced enum, which is what the previous version of this site did and why it was deleted."
    },
    {
      "field": "toxicity",
      "status": "not_published",
      "reason": "Not published. The widely copied list is a copyrighted compilation restricted to noncommercial use, and we will not scrape it. A sourced toxicity dataset is being built from primary literature and is not ready."
    },
    {
      "field": "botanical_review",
      "status": "not_sourced",
      "reason": "No page on this site has been reviewed by a named botanist. The determination label says so, on every page."
    },
    {
      "field": "native_range",
      "status": "not_sourced",
      "reason": "Kew's World Checklist of Vascular Plants does not cover this taxon."
    },
    {
      "field": "hardiness_zone",
      "status": "not_published",
      "reason": "Not published for any species. A USDA hardiness zone is the average annual EXTREME minimum temperature, and we hold the mean daily minimum of the coldest month, which is a different quantity and typically about 16 C warmer. Presenting one as the other would place a plant several zones too warm, so we do not publish a zone at all."
    },
    {
      "field": "phenology",
      "status": "not_sourced",
      "reason": "No dated flowering annotations found for this taxon."
    }
  ],
  "license": {
    "data": "CC BY 4.0",
    "attribution": "BotanAI, https://botanai.app/plants/geheebia-tophacea/",
    "note": "This document is licensed CC BY 4.0. The PHOTOGRAPHS it links are licensed individually and separately: each carries its own license and license_url, and those terms govern. Underlying taxonomy is CC0 (World Flora Online); native range is CC BY 3.0 (Kew)."
  }
}