Athyrium spinulosum(Maxim.) Milde

WFO wfo-0001116125 Accepted WFO 2026-06 7 photographs CC BY

Plate 1 figs. a–g · 3 observations

This species has been photographed under an open licence only 3 times, so some figures below are different views of the same plant, taken on the same day, rather than different individuals. They are usually different parts of it: the leaf, the flower, the bark.

Athyrium spinulosum, photographed by Nina Filippova
fig. a Nina Filippova, CC BY 4.0 / 2021-09-03 / obs. 157728852
Athyrium spinulosum, photographed by Nina Filippova
fig. b Nina Filippova, CC BY 4.0 / obs. 153672155
Athyrium spinulosum, photographed by Oleg Kosterin
fig. c Oleg Kosterin, CC BY 4.0 / obs. 98731200
Athyrium spinulosum, photographed by Nina Filippova
fig. d Nina Filippova, CC BY 4.0 / obs. 157728888
Athyrium spinulosum, photographed by Nina Filippova
fig. e Nina Filippova, CC BY 4.0 / obs. 153672078
Athyrium spinulosum, photographed by Nina Filippova
fig. f Nina Filippova, CC BY 4.0 / obs. 153672114
Athyrium spinulosum, photographed by Nina Filippova
fig. g Nina Filippova, CC BY 4.0 / obs. 153672040

Every figure is a research-grade observation under CC0, CC BY or CC BY-SA, rehosted with the photographer’s name, the licence and the observation it came from. Photographs under a NonCommercial licence are excluded from this site and are never stored, which costs us a great many pictures and is not negotiable.

The specimen a real sheet, in a real collection

Herbarium
The New York Botanical Garden
Accession
04058454
Filed as
Athyrium spinulosum (Maxim.) Milde
Det. by
not recorded on this sheet
Collected
E. Faber
Origin
CN
The sheet
View the digitised specimen (CC BY 4.0)

A real pressed plant, in a real collection, under the accession number above. Not an illustration of one. The holding institution does not serve this sheet’s image to third parties, so there is no photograph here. The record is real and the link goes to it. Where we hold no openly licensed sheet for a species this section is simply absent, and where a sheet never recorded who determined it, that field stays empty rather than being filled in. Roughly half of all herbarium sheets never recorded a determiner, which is ordinary.

Native range 15 botanical countries

Regions where Athyrium spinulosum is native: Amur, China North-Central, China South-Central, China Southeast, Inner Mongolia, Japan, Khabarovsk, Korea, Manchuria, Primorye, Qinghai, Sakhalin, Tibet, East Himalaya, Nepal AmurChina North-CentralChina South-CentralChina SoutheastInner MongoliaJapanKhabarovskManchuriaPrimoryeQinghaiSakhalinTibetEast HimalayaNepal Korea
Native distribution of Athyrium spinulosum, after Kew’s World Checklist of Vascular Plants. Introduced, extinct and doubtful records are excluded, so this is where the plant is from, not everywhere it now grows. Regions too small to draw at this scale are marked with a dot.
RegionTDWG codeContinent
Amur AMU ASIA-TEMPERATE
China North-Central CHN
China South-Central CHC
China Southeast CHS
Inner Mongolia CHI
Japan JAP
Khabarovsk KHA
Korea KOR
Manchuria CHM
Primorye PRM
Qinghai CHQ
Sakhalin SAK
Tibet CHT
East Himalaya EHM ASIA-TROPICAL
Nepal NEP

Region boundaries approximated from Natural Earth (public domain) and mapped to TDWG World Geographical Scheme for Recording Plant Distributions (WGSRPD) level-3 botanical countries (Brummitt 2001). Indicative, not the official WGSRPD geometry.

Where it actually grows measured, from 73 observations

Condition 5th percentile Median 95th percentile
Coldest month, mean daily low -22.2 °C -16.3 °C -12.6 °C
Warmest month, mean daily high 20.8 °C 22.3 °C 24.2 °C
Annual rainfall 638 mm 822 mm 995 mm
Rainfall in the driest quarter 37 mm 46 mm 71 mm

It is found where winters are severely cold. This is not care advice and it is not a forecast. It is a measurement: we sampled the climate at every one of the 73 research-grade observations of Athyrium spinulosum that carry a coordinate, and this is the range those places actually span. The 5th and 95th percentiles are used rather than the minimum and maximum, because a single cultivated specimen in a heated conservatory should not widen a tropical plant's range to the Arctic.

This is not a hardiness zone. A USDA zone is the average annual extreme minimum temperature. The figure above is the mean daily minimum of the coldest month, which is a different quantity and is typically far warmer. Reading one as the other would place a plant several zones too warm, so we do not publish a hardiness zone, because we do not have one.

Also published as 23 synonyms

A synonym is not an error. It is a record of botanists disagreeing, in print, about where this plant belongs. Each of these was somebody’s considered answer.

  • Asplenium spinulosum (Maxim.) Miq.
  • Asplenium spinulosum C.B.Clarke
  • Asplenium spinulosum (Maxim.) Baker
  • Asplenium spinulosum var. subtriangulare (Hook.) C.B.Clarke
  • Asplenium subtriangulare Hook.
  • Athyrium hookerianum T.Moore
  • Athyrium longipes Christ
  • Athyrium sikkimense (Bir) Á.Löve & D.Löve
  • Athyrium spinulosum var. subtriangulare (Hook.) C.Chr.
  • Athyrium subtriangulare (Hook.) Bedd.
  • Athyrium subtriangulare var. sikkimense Bir
  • Cystopteris spinulosa Maxim.
  • Pseudocystopteris decipiens Ching & S.K.Wu
  • Pseudocystopteris longipes (Christ) Ching
  • Pseudocystopteris purpurascens Ching & S.K.Wu
  • Pseudocystopteris reflexipinnula Ching & S.K.Wu
  • Pseudocystopteris remota Ching
  • Pseudocystopteris sikkimensis (Bir) Ching
  • Pseudocystopteris sparsa Ching & S.K.Wu
  • Pseudocystopteris spinulosa (Maxim.) Ching
  • Pseudocystopteris spinulosa var. taipaishanensis Ching
  • Pseudocystopteris subtriangularis (Hook.) Ching
  • Pseudocystopteris tibetica Ching

Sourcesevery claim on this page

  1. World Flora Online Plant List. accepted name, authority, classification. CC0. Retrieved 2026-07-12.
  2. iNaturalist. photographs and flowering annotations, CC0 / CC BY / CC BY-SA only. per photograph. Retrieved 2026-06-27.
  3. Kew, World Checklist of Vascular Plants (WCVP v16). native distribution by TDWG level-3 botanical country, and life form. CC BY 3.0. Retrieved 2026-06-04.

We publish what we can source and we say so when we cannot. This page has no care advice and no toxicity claim, because we do not yet have those from a source we can cite.